5 Ma and increased its abundance thereafter. The increased magnitude of fluctuations in the relative abundances of Bulimina aculeata, species diversity and the percentage of total infaunal taxa during the Pleistocene reflects significant variations in the trophic level due to changing surface water productivity, possibly

in response to glacial/interglacial changes. The relatively larger fluctuations in diversity values during the Pleistocene can be explained as a faunal response to the glacial and interglacial cycles with the varying eastern Afatinib concentration Asian monsoon regime ( An 2000) and thus changing trophic conditions for the benthic foraminifera ( Rai & Singh 2001). The disappearance of Stilostomella lepidula during the middle Pleistocene (∼ 0.7 Ma) almost coincides with the so-called global ‘Stilostomella’ extinction ( Schönfeld, 1996, Hayward, 2001 and Hayward, 2002), whereas this species has also been recorded with rare and sporadic occurrences in the recent sediment of the Indian Ocean and other regions (see Rai & Singh 2004). The dominant occurrence Ribociclib of the B. aculeata assemblage during the last ∼ 0.7 Ma suggests that the increase in upwelling and surface water productivity is possibly responsible for the sudden decline of oligotrophic Stilostomella

and the almost complete absence of the C. lobatulus assemblage in the Indian Ocean. Gupta & Thomas (1999) also suggested that the decline and loss of this group was due mainly to intensified cooling combined with increased upwelling and surface water productivity,

and in places the increased strength of tropical monsoons. The closing of the Indonesian seaway was responsible for several palaeoceanographic changes in the eastern Indian Ocean. The final closure of the Indonesian seaway at about 4–3 Ma changed the source of the Indonesian Throughflow (ITF) from warm south Pacific to cold north Pacific waters, which resulted in the breakup of permanent El Niño-like conditions. These changes reduced the warm thermocline water Sitaxentan flow into the eastern Indian Ocean and also started the development of upwelling-led higher surface water productivity systems in this region. The flow of northern cold Pacific waters into the Indian Ocean may have lowered SSTs in upwelling regions, which caused the cooling of northern America through teleconnections and also initiated the late Pliocene glaciations in the Northern Hemisphere. The changing strength of the southward-flowing warm Leeuwin Current (LC) and the northward-flowing cold Western Australian Current (WAC) in response to glacial/interglacial cycles may have played an important role in the oceanographic setting of this area during the Pleistocene.